First episode (2005): "A Hard Day's Night" (Season 1, Episode 1)

Last episode (2009): "Now or Never" (Season 5, Episode 24)

What he's up to now (2020): After Grey's, T.R. appeared in several episodes of The Good Wife, and he rejoined the ShondaLand family in the short-lived series The Catch. Most recently, he starred in Genius: Einstein and Genius: Picasso. He'll appear in the upcoming limited series A Higher Loyalty opposite Jeff Daniels, Holly Hunter, and Brendan Gleeson, among others.